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14-2                                Feeds and Feeding                                CH 14]




                                Swine, horses and mules, are monogastric animals. As the word monogastric suggests, this type
                            of digestive system consists of one (“mono”) stomach chamber (“gastric”). The process of digestion
                            begins with the mouth and the intake of food.

             Monogastric:       Poultry have a boney beak and no teeth, therefore they do not chew (masticate) their food.
             Single-chambered   Poultry feed passes into the birds crop. A chicken's crop has a specific function which is to sort food
             Stomach.       out like a holding tank. When the food is ready and sorted out in the crop, it passes into one of the two

                            gastric chambers that a chicken has, which are the proventriculus or the gizzard. The “easy” to digest
             Proventriculus: A   food goes to the proventriculus where it's broken down by hydrochloric acid as well as digestive
             standard part of   enzymes. Food that is harder to digest and which needs to be “mechanically” broken down passes to
             avian anatomy,   the gizzard to be processed - this is where muscular contractions combined with some small stones
             and is a rod   (grit) are needed to process food that is ingested.
             shaped organ,
             located between    As there are thousands of feedstuffs used to formulate animal rations, a discussion of feedstuffs
             the esophagus and   could be made on the basis of groups or type of feedstuffs by their common characteristics. The
             the gizzard of most   National Research Council (NRC) in 1972 established such groups. It is important to realize that the
             birds. It is generally   classification of feedstuffs into categories is imprecise because most feedstuffs are complex multiple
             a glandular part of
             the stomach that   nutrient sources. For example, the same feedstuff may legitimately be considered either an energy
             may store and/or   feed or a protein supplement, or even a forage, such as Alfalfa, which is a roughage, with a 15.3%
             commence       protein content, a 1.9% fat content and a 28.6% fiber content. Upon being digested, the energy used to
             digestion of food   produce cell development, maintenance, and growth come predominantly from the protein as
             before it
             progresses to the   digestible protein and fat.
             gizzard.           Protein provides 4 calories of energy for every gram of protein consumed, and is the same
                            amount derived from carbohydrates; fats deliver 9 calories per gram. Extra calories consumed are
                            stored as fat because it's such a concentrated source of energy. When a body requires energy, it first
             Gizzard:  The hind
             part of the    uses glucose from carbohydrates, then fatty acids. As long as enough calories are consumed from
             stomach,       other sources, protein is not turned into energy.
             especially modified
             for grinding food.
             The gizzard has a
             thick muscular wall   Utilization of Energy            Figure 14.1
             and may contain
             small stones, or       GROSS ENERGY
             gastroliths, that
             function in the                                                Additional energy is needed for gains of body tissue,
             mechanical     FECAL ENERGY                                    any increase in energy stores, growth of the fetus
             breakdown of                                                   during pregnancy, production of milk during lactation,
             seeds and other                                                and energy losses associated with synthesis/deposition
             foods.                         DIGESTIBLE ENERGY               of new tissue or milk.
                                              (COMPARABLE TO TDN)
                                                                            TDN: Total Digestible Nutrients

                                                                            Source: Adapted from Warwick and Baines (2000) and
                                                                            Livesey.
                                      URINARY AND
                                    COMBUSTIBLE GAS
                                        ENERGY

                                                      METABOLIZABLE ENERGY


                                                Heat Increment

                                                                     NET  ENERGY


                                                     NET ENERGY MAINTENANCE      NET ENERGY
                                                       Expelled as gas through
                                                      respiration and perspiration   PRODUCTION




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